This
is one of the smallest of the grain-infesting beetles, but one of the
most important. Originally native to the tropics, it has spread through
commerce to all parts of the world. It is most prominent in the United
States, southern Canada, Argentina, India, and Australia. In the United
States, it is particularly widespread in the Gulf states. Both adults
and larvae feed within the interiors of nearly all grains, including rice,
and the kernels are reduced to mere shells, hence it is a hidden infestation
until the adults are apparent.
Description:
The
adult is about 3 mm long, polished dark brown or black, and has a somewhat
roughened surface. In common with other bostrichids, which are principally
woodborers, Rhyzopertha dominica is almost cylindrical,
and the head is vertically deflexed under the thorax so that it cannot
be seen from above. Likewise, this insect, though small, has powerful
jaws with which it can bore directly into wood . Wood may have been its
original food. It can eat its way into wooden and paper boxes, and may
destroy book bindings.

Photograph
courtesy of Jim Kalisch University of Nebraska
Life
Cycle:
The
female lays 300 to 500 eggs, singly or in clusters, in the loose grain.
They hatch in a few days. The larvae moult 2 to 4 times. They may feed
on the flour produced by the boring of the adults, or may bore directly
into kernels that have been slightly damaged. They complete their growth
within the grain, transform to white pupae, and the adults cut their way
out. The life cycle takes only a month or two, depending on the temperature.
The adult lesser grain borers chews
grain voraciously causing damage which may facilitate infestation by a
secondary pest. It is a strong flyer and may rapidly migrate from infested
grain to begin new infestations elsewhere.
Factors
Favoring Stored Grain Insects:
Temperature
is a crucial environmental factor that influences the development of insects.
There is always a minimum, optimum and a maximum range of temperature
in which insects can survive. The most favorable temperature for most
stored grain insects is about 80°F. Above 95°F or below 60°F, reproduction
and survival is greatly reduced. Insects differ in their tolerance to
either low or high temperatures. Most stored product pests would follow
the same pattern of survival under a different range of temperatures.
As temperature approaches zero, insect development, activity and movement
decline to a minimum. Gradual increase in temperature will increase insect
activity up to a certain range that differs among different species. Further
increase in temperature above the optimum range will lead to increase
in insect mortality and crashing of the population.
The
most favorable moisture range for stored grain insects is 12 to 15% (optimum
14 to 15%). The lowest moisture content for any survival and reproduction
to occur is about nine percent. However, as temperature increases insects
can reproduce in grain with a lower moisture content, and when moisture
increases they can reproduce at lower temperatures. The life processes
of the insects can increase the moisture content of "dry" grain.
Insect
infestations are more likely in "dirty" grain than in clean grain. The
presence of grain dust, cracked kernels, and dockage permits some grain
insects to survive and reproduce at low moisture levels.

Photograph
courtesy of Jim Kalisch University of Nebraska
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